TY - JOUR TI - Cryo-EM structures of amyloid-beta filaments with the Arctic mutation (E22G) from human and mouse brains EP - 333 SP - 325 AV - public KW - Science & Technology KW - Life Sciences & Biomedicine KW - Clinical Neurology KW - Neurosciences KW - Pathology KW - Neurosciences & Neurology KW - Alzheimer's disease KW - Amyloid-beta KW - Arctic mutation KW - Electron cryo-microscopy KW - Mouse App(NL-G-F) knock-in line KW - Tau KW - HEREDITARY CEREBRAL-HEMORRHAGE KW - BEAM-INDUCED MOTION KW - ALZHEIMERS-DISEASE KW - GENE KW - PEPTIDES VL - 145 N2 - The Arctic mutation, encoding E693G in the amyloid precursor protein (APP) gene [E22G in amyloid-? (A?)], causes dominantly inherited Alzheimer?s disease. Here, we report the high-resolution cryo-EM structures of A? filaments from the frontal cortex of a previously described case (A?PParc1) with the Arctic mutation. Most filaments consist of two pairs of non-identical protofilaments that comprise residues V12?V40 (human Arctic fold A) and E11?G37 (human Arctic fold B). They have a substructure (residues F20?G37) in common with the folds of type I and type II A?42. When compared to the structures of wild-type A?42 filaments, there are subtle conformational changes in the human Arctic folds, because of the lack of a side chain at G22, which may strengthen hydrogen bonding between mutant A? molecules and promote filament formation. A minority of A?42 filaments of type II was also present, as were tau paired helical filaments. In addition, we report the cryo-EM structures of A? filaments with the Arctic mutation from mouse knock-in line AppNL?G?F. Most filaments are made of two identical mutant protofilaments that extend from D1 to G37 (AppNL?G?F murine Arctic fold). In a minority of filaments, two dimeric folds pack against each other in an anti-parallel fashion. The AppNL?G?F murine Arctic fold differs from the human Arctic folds, but shares some substructure. JF - Acta Neuropathologica UR - https://doi.org/10.1007/s00401-022-02533-1 PB - SPRINGER N1 - Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons licence, and indicate if changes were made. The images or other third party material in this article are included in the article's Creative Commons licence, unless indicated otherwise in a credit line to the material. If material is not included in the article's Creative Commons licence and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this licence, visit http://creativecommons.org/licenses/by/4.0/. IS - 3 Y1 - 2023/01/07/ ID - discovery10168368 SN - 0001-6322 A1 - Yang, Yang A1 - Zhang, Wenjuan A1 - Murzin, Alexey G A1 - Schweighauser, Manuel A1 - Huang, Melissa A1 - Lovestam, Sofia A1 - Peak-Chew, Sew Y A1 - Saito, Takashi A1 - Saido, Takaomi C A1 - Macdonald, Jennifer A1 - Lavenir, Isabelle A1 - Ghetti, Bernardino A1 - Graff, Caroline A1 - Kumar, Amit A1 - Nordberg, Agneta A1 - Goedert, Michel A1 - Scheres, Sjors HW ER -