eprintid: 10189805 rev_number: 13 eprint_status: archive userid: 699 dir: disk0/10/18/98/05 datestamp: 2024-03-27 08:55:32 lastmod: 2024-03-27 08:55:32 status_changed: 2024-03-27 08:55:32 type: article metadata_visibility: show sword_depositor: 699 creators_name: Van der Ham, Stijn creators_name: Agudo-Canalejo, Jaime creators_name: Vutukuri, Hanumantha Rao title: Role of Shape in Particle-Lipid Membrane Interactions: From Surfing to Full Engulfment ispublished: inpress divisions: UCL divisions: B04 divisions: C06 divisions: F60 keywords: Vesicles; Lipid membranes; Anisotropic particles; Wrapping; Passive engulfment; Cellular particle uptake note: This article is licensed under CC-BY 4.0, https://creativecommons.org/licenses/by/4.0/. abstract: Understanding and manipulating the interactions between foreign bodies and cell membranes during endo- and phagocytosis is of paramount importance, not only for the fate of living cells but also for numerous biomedical applications. This study aims to elucidate the role of variables such as anisotropic particle shape, curvature, orientation, membrane tension, and adhesive strength in this essential process using a minimal experimental biomimetic system comprising giant unilamellar vesicles and rod-like particles with different curvatures and aspect ratios. We find that the particle wrapping process is dictated by the balance between the elastic free energy penalty and adhesion free energy gain, leading to two distinct engulfment pathways, tip-first and side-first, emphasizing the significance of the particle orientation in determining the pathway. Moreover, our experimental results are consistent with theoretical predictions in a state diagram, showcasing how to control the wrapping pathway from surfing to partial to complete wrapping by the interplay between membrane tension and adhesive strength. At moderate particle concentrations, we observed the formation of rod clusters, which exhibited cooperative and sequential wrapping. Our study contributes to a comprehensive understanding of the mechanistic intricacies of endocytosis by highlighting how the interplay between the anisotropic particle shape, curvature, orientation, membrane tension, and adhesive strength can influence the engulfment pathway. date: 2024-03-21 date_type: published publisher: American Chemical Society (ACS) official_url: http://dx.doi.org/10.1021/acsnano.3c11106 oa_status: green full_text_type: pub language: eng primo: open primo_central: open_green verified: verified_manual elements_id: 2262511 doi: 10.1021/acsnano.3c11106 medium: Print-Electronic lyricists_name: Agudo-Canalejo, Jaime lyricists_id: JAGUD28 actors_name: Agudo-Canalejo, Jaime actors_id: JAGUD28 actors_role: owner funding_acknowledgements: [Dutch Research Council (NWO)] full_text_status: public publication: ACS Nano event_location: United States issn: 1936-0851 citation: Van der Ham, Stijn; Agudo-Canalejo, Jaime; Vutukuri, Hanumantha Rao; (2024) Role of Shape in Particle-Lipid Membrane Interactions: From Surfing to Full Engulfment. ACS Nano 10.1021/acsnano.3c11106 <https://doi.org/10.1021/acsnano.3c11106>. (In press). Green open access document_url: https://discovery-pp.ucl.ac.uk/id/eprint/10189805/1/Agudo-Canalejo_van-der-ham-et-al-2024-role-of-shape-in-particle-lipid-membrane-interactions-from-surfing-to-full-engulfment.pdf